MSc(Res) Marine Biology

The Master of Science by Research degree in Marine Biology is a 12-month, research only degree, in which the candidate will undertake a supervised research project in the area of Marine Biology, in the School of Biology, University of St Andrews.

The candidate will be based in the interdisciplinary Scottish Oceans Institute (SOI), based at East Sands, St Andrews. The SOI is home to the world famous Sea Mammal Research Unit (SMRU), and hosts a wide-range of researchers with interests ranging from the deep oceans to the coasts, and from the people who use and interact with the sea, to the biological and physical processes that make the oceans function.

SOI supports four research themes:

  • Global Change and Planetary Evolution
  • Ecology, Fisheries and Resource management
  • Developmental and Evolutionary Genomics
  • The Sea Mammal Research Unit.

SOI brings together behavioural and population ecologists, evolutionary biologists, geneticists, conservationists, mathematicians and environmental scientists to contribute towards understanding and managing the future of our oceans and the organisms, big and small, that live within them. Further details about the SOI.

Candidates may approach potential supervisors in the SOI directly or via advertised projects listed below.



Evolutionary developmental biology

Supervisor: Dr David E. K. Ferrier

Research area(s): Evolutionary developmental biology

Research description: We seek to understand how the diversity of animal forms have evolved via changes to their development, usually taking the homeobox genes of the Hox/ParaHox and related clusters as a starting point. A variety of invertebrate species are studied, with the aim of focusing on major transitions in animal evolution, including the origins of the animal kingdom, the origin of the bilaterally symmetrical animals (bilaterians) and the origin of chordates and vertebrates.

Relevant references: See

Subject area(s): Evolutionary biology, Developmental Biology

Keywords: homeobox genes, amphioxus, Ciona, polychaetes

Evolutionary biology, developmental biology, cell biology, evo-devo, regeneration biology, genomics

Supervisor: Dr Ildiko Somorjai

Research area(s): Evolutionary biology, developmental biology, cell biology, evo-devo, regeneration biology, genomics

Research description: Have you ever wondered why some animals regenerate well, and humans do not? Are you interested in how new genes are born, and what generates diversity in animal body forms? The Somorjai lab addresses these problems from evolutionary, developmental and cell biological perspectives. We predominantly use the marine invertebrate chordate “amphioxus” due to its genetic and anatomical similarly to simple vertebrates. We also work on flatworms, which have amazing regenerative powers and multipotent stem cells. The project will depend on the student’s interests and background, but could include gene expression analyses, embryology, immunohistochemistry, confocal microscopy, genomics, and phylogenetic analyses.

Relevant References:

Bertrand S, Escriva H. Evolutionary crossroads in developmental biology: amphioxus. Development. 2011 Nov;138(22):4819-30.

Somorjai IM, Somorjai RL, Garcia-Fernàndez J, Escrivà H. Vertebrate-like regeneration in the invertebrate chordate amphioxus. Proc Natl Acad Sci U S A. 2012 109(2):517-22.

Dailey, SC, Planas, RF, Espier, AR, Garcia-Fernandez, J & Somorjai, IML Asymmetric distribution of pl10 and bruno2, new members of a conserved core of early germline determinants in cephalochordates. Frontiers in Ecology and Evolution. 2016. 3, 156.

Subject area(s): Evolutionary Biology, Developmental Biology

Keywords: Regeneration, Development, Evo-devo, Amphioxus


Entry requirements and selection process

An undergraduate Honours degree at 2:1 level or better in biological or environmental sciences. Students from backgrounds such as mathematics and the physical or chemical sciences may be accepted under exceptional circumstances.

If you studied for your first degree outside of the UK, please see the international entry requirements

For non-native English speakers, please see the English language requirements

Applicants will be short-listed by the project supervisor, and subject to interview by the project supervisor and an additional member of the Biology Postgraduate Committee.


For details of postgraduate tuition fees relevant to our research degrees including the MSc(Res), please visit:

Progression and assessment

Students in the MSc(Res) will be assigned an Internal Examiner (IE) and PG Tutor by the School. There will be a progress review meeting at three months to monitor and evaluate student progression, convened by the IE, with the student and Tutor in attendance. This meeting will be guided by a brief supervisor report and will be based on oral examination with no requirement for a written submission by the student.

The degree requires submission and examination of a dissertation at the end of the one-year program. As per 2016-2017 Senate Regulations (page 9), this thesis will consist of up to 30,000 words. The thesis will be evaluated by the IE and an External Examiner appointed at time of submission. Evaluation will be based on the written submission; there is no requirement for a viva.

Skills Training

In addition to the project-specific training that you will receive during your degree, Msc(Res) students will also have access to a wide range of training in transferable skills through the award-winning University of St Andrews GradSkills program, run by our Professional Development Unit CAPOD.

Specific post-graduate programs run within the School of Biology may also offer additional training, for instance in statistical, bioinformatics or molecular techniques.


Students may apply for placement in advertised projects or contact potential supervisors directly. We strongly recommend that potential candidates make contact with a potential supervisor before applying.  See links on this page.

Biology has two dates for admission to this degree: September and January each year.

If you have decided that you would like to make a formal application to study for an MSc(Res) at St Andrews, please complete an application using the online system.

Note: If you are self-funded and interested in working with a supervisor who does not currently have a project listed, please contact that person directly: supervisors’ email addresses may be found using the links on this page.