I’m very happy to support applications for research fellowships, for example Royal Society Newton International Fellowships, Marie Skłodowska-Curie Research Fellowships, Royal Commission for the Exhibition of 1851 Research Fellowships, FCT Post-doctoral Fellowships, etc. I’ve had success in obtaining  research fellowships for myself throughout my career, and have recently welcomed two early-career researchers to my group on such funding schemes. So do get in touch if you are interested in bringing a fellowship my research group.


I am advertising a PhD project on

Explaining human handedness: the role of parental genes

which will be co-supervised by me and Dr Silvia Paracchini, jointly between the Schools of Biology and Medicine at the University of St Andrews. This is a fully-funded PhD studentship, with tuition fees (at both Home and Overseas rates), research costs and living allowance covered by the European Research Council and the School of Biology, University of St Andrews. Full details are given here. The deadline for applications is 5 Jan 2021.

A further PhD opportunity is also available in the research group of my collaborator Dr Laura Ross, at the University of Edinburgh. I will be co-supervising this project.

Sex, males & hermaphrodites in the scale insect Icerya purchasi

Full details of this PhD position are given here. The deadline for applications is 6 Jan 2021.

Image from Wikimedia Commons.


Even when I’m not explicitly advertising a position, there is the possibility of us applying for funding for your project. For example: graduates of Scottish universities who wish to undertake their PhD here at St Andrews may be eligible for Carnegie funding; St Andrews offers 30+ PhD studentships every year to Chinese nationals in conjunction with CSC; and I’ve had several Portuguese students undertake their PhD studies in my lab with the support of FCT. Also, St Andrews now offers a number of St Leonard’s PhD Scholarships for PhD students engaged in interdisciplinary work linking two Schools within the University and also for PhD students jointly enrolled at St Andrews and another European university. So there are lots of options for PhD funding.

Alternatively, there is also the possibility of pursuing a one-year research degree in my research group, in the form of a MSc(Res). Do get in touch if you think you may be interested in this option.


I offer theoretical projects for honours students, typically but not exclusively on the topic of social evolution.

School & College

I offer 4-6 week theoretical projects as part of the Nuffield Research Placements scheme.