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Information for current students at the School of Biology

Biology Online Module Handbook

BL3316 Co evolution living together

(BL3316 online module handbook version 87)

cover image

image: Hummingbird pollinating flower

This module considers coevolution: how organisms influence the evolution of other organisms around them. Some of the most exquisite and remarkable adaptations in the natural world are the products of coevolution, from the elaborate displays of birds-of-paradise to the sexual mimicry of orchids. We will consider coevolution within a species, focusing in particular on the interactions between males and females, and also coevolution between species, including animal-plant interactions, host-parasite interactions, and predator-prey interactions. Topics to be considered in detail will be sexual selection, pollination biology, herbivory, prey defences, the evolution of virulence, and the ecology of coevolution, focusing in particular on tritrophic interactions between animals, plants and fungi. Finally, we will consider broader themes in coevolution, including the geographic mosaic theory of coevolution and applied aspects of coevolution.*

Credits: 20*

Semester: 2

Module Organiser
Dr Verena Dietrich-Bischoff
vdb@st-andrews.ac.uk
01334 464812
Pre-requisite Modules:
Before taking this module you must pass BL2304 or pass BL2307 or pass BL2312*
Anti-requisite Modules:
*
Post-requisite Modules:
*
Additional Module Information:


Biology Teaching Office:
Level 2, Biomolecular Sciences Building, University of St Andrews, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk      Phone: +44 (0)1334 46 3602 or 3566

BL3316: Timetable

Legend (not all modules have every event type):
lecture
tutorial
workshop
practical
other

Semester 2: Week 1

DATE & TIMEVENUESTAFFEVENT
Monday
15-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L1: Overview of the module and introduction to coevolution
2023-4_BL3316_L1
Tuesday
16-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L2: The geographic mosaic theory of coevolution
2023-4_BL3316_L2
Wednesday
17-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Tutorial T1: Paper discussion (Synthesis of coevolution across levels of biological organisation)
2023-4_BL3316_T1

Semester 2: Week 2

DATE & TIMEVENUESTAFFEVENT
Monday
22-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L3: Pollination
2023-4_BL3316_L3
Tuesday
23-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L4: Pollination and coevolution
2023-4_BL3316_L4
Wednesday
24-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L5: Seed dispersal
2023-4_BL3316_L5

Semester 2: Week 3

DATE & TIMEVENUESTAFFEVENT
Monday
29-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L6: Coevolution of fruit colouration
2023-4_BL3316_L6
Tuesday
30-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L7: Coevolution of other fruit traits
2023-4_BL3316_L7
Wednesday
31-01-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Tutorial T2: Discussion (How do coevolutionary interactions differ between pollination and seed dispersal?)
2023-4_BL3316_T2

Semester 2: Week 4

DATE & TIMEVENUESTAFFEVENT
Monday
05-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Harry Watkins
-
Lecture L8: The role of botanic gardens in the late Anthropocene
2023-4_BL3316_L8
Tuesday
06-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Harry Watkins
-
Lecture L9: Green infrastructure: harnessing urban ecologies as a frontier of evolution
2023-4_BL3316_L9
Wednesday
07-02-2024
13:00 to 17:00
Other
St Andrews Botanic Garden
Dr Harry Watkins
-
Practical P1: Exploring the leaf economic spectrum in Northeast Fife
2023-4_BL3316_P1

Semester 2: Week 5

DATE & TIMEVENUESTAFFEVENT
Monday
12-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L10: Ant-plant interactions 1
2023-4_BL3316_L10
Tuesday
13-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L11: Ant-plant interactions 2
2023-4_BL3316_L11
Wednesday
14-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L12: Figs and fig wasps
2023-4_BL3316_L12

Semester 2: Week 6

DATE & TIMEVENUESTAFFEVENT
Monday
19-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Miguel Barbosa
-
Lecture L13: Coevolution on the coral reef
2023-4_BL3316_L13
Tuesday
20-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L14: Fish cleaning mutualisms
2023-4_BL3316_L14
Wednesday
21-02-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Tutorial T3: Discussion (Cleaning mutualisms) and introduction to assessment
2023-4_BL3316_T3

Spring Break: 26-Feb-2024 to 01-Mar-2024

Semester 2: Week 7

DATE & TIMEVENUESTAFFEVENT
Monday
04-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L15: Concepts in predator-prey coevolution
2023-4_BL3316_L15
Tuesday
05-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L16: Coevolution in echolocating predators and their prey
2023-4_BL3316_L16
Wednesday
06-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Lecture L17: Carnivorous plants
2023-4_BL3316_L17

Semester 2: Week 8

DATE & TIMEVENUESTAFFEVENT
Monday
11-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L18: Avian brood parasitism
2023-4_BL3316_L18
Tuesday
12-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L19: Insect social parasitism
2023-4_BL3316_L19
Tuesday
12-03-2024
14:00 to 17:00
Biomolecular Sciences Building
BMS 205a
Dr Verena Dietrich-Bischoff
-
Practical P2: Exploring egg rejection defences against brood parasites
2023-4_BL3316_P2
Wednesday
13-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Tutorial T4: Paper discussion (Brood parasitism in fish)
2023-4_BL3316_T4

Semester 2: Week 9

DATE & TIMEVENUESTAFFEVENT
Monday
18-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Nathan Bailey
-
Lecture L20: Host-parasite coevolution I
2023-4_BL3316_L20
Tuesday
19-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Nathan Bailey
-
Lecture L21: Host-parasite coevolution II
2023-4_BL3316_L21
Wednesday
20-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Graeme Ruxton
-
Tutorial T5: Paper discussion (Birds using cigarette buds to control nest parasites)
2023-4_BL3316_T5

Semester 2: Week 10

DATE & TIMEVENUESTAFFEVENT
Tuesday
26-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L22: Sexual selection as coevolution
2023-4_BL3316_L22
Wednesday
27-03-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof Mike Ritchie
-
Lecture L23: Coevolution in the genome
2023-4_BL3316_L23

Semester 2: Week 11

DATE & TIMEVENUESTAFFEVENT
Monday
01-04-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L24: Sexual conflict
2023-4_BL3316_L24
Tuesday
02-04-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Dr Verena Dietrich-Bischoff
-
Lecture L25: Effects of climate change and other anthropogenic stressors on coevolutionary relationships
2023-4_BL3316_L25
Wednesday
03-04-2024
13:00 to 14:00
Bute Building
Lecture Theatre A
Prof John Jones
-
Lecture L26: Plant-nematode interactions and their implications for agriculture
2023-4_BL3316_L26
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3316: Assessment

3-hour Written Examination = 50%, Coursework = 50%*

The following related information applies to all Biology modules:

School of Biology Marking Criteria: See JH booklet info (st-andrews.ac.uk) 
Late submission of continuous assessment work:

All late submissions of coursework that do not require electronic submission should be made via the Biology Teaching Office, Level 2, BMS Building, North Haugh.
Exam details: See School of Biology UG Handbook JH booklet info (st-andrews.ac.uk)  : All Biology exams will be conducted online for 2022-23.
Exam timetable: See Timetables - Exams - University of St Andrews (st-andrews.ac.uk) 
Expected attendance: See JH booklet info (st-andrews.ac.uk)  for detailed attendance requirements.
Good Academic Practice & Avoiding Academic Misconduct: See JH booklet info (st-andrews.ac.uk)
University Student Handbook: University Student Handbook
School and University regulations in the School and University Undergraduate Handbook relating to absence reporting, penalties and rules for late submission of work, extensions for coursework, return of coursework, S-coding, good academic practice and Academic Alerts.: JH booklet info (st-andrews.ac.uk)
University Student Handbook
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Who to ask

(Information in this section applies to all Biology Modules)

Before contacting staff, students should check the content of the Biology Undergraduate Handbook, the module handbook and specific task instructions.

Questions about Contact
General teaching matters Biology Teaching Office ( bioteach@st-andrews.ac.uk )
Rescheduled or cancelled events Check your University email
Lecture or practical content The lecturer who presented the material
Completing assessed practical assignments The lecturer who set the assignment
Completing assessments Module Organiser (Dr Verena Dietrich-Bischoff vdb@st-andrews.ac.uk)
Marking on continuous assessment The Demonstrator or Module Organiser (Dr Verena Dietrich-Bischoff vdb@st-andrews.ac.uk)
Marking on exams Module Organiser (Dr Verena Dietrich-Bischoff vdb@st-andrews.ac.uk)
Rearranging practical days
Module Organiser (Dr Verena Dietrich-Bischoff vdb@st-andrews.ac.uk)
Absence and/or extensions Module Organiser (Dr Verena Dietrich-Bischoff vdb@st-andrews.ac.uk)

and the Biology Teaching Office ( bioteach@st-andrews.ac.uk )

Difficulties with academic progress which impact more than one module:

Year Coordinator

See School of Biology UG Handbook for list:
JH booklet info (st-andrews.ac.uk)

Overall performance, progress or future directions:

Advisor of Studies

Disability:

Disability Coordinator ( biodisabilities@st-andrews.ac.uk )

For advice and support on any issue e.g. academic, financial, international, personal or health matters, or if you are unsure of who to go to for help:

Advice & Support Centre

Address: 79 North Street, St Andrews

Email: theasc@st-andrews.ac.uk

Web: https://www.standrews.ac.uk/ask-a-question/

Tel: 01334 462020

University assistance with urgent matters out of office hours:

Tel: 01334 476161

Web: https://www.st-andrews.ac.uk/students/advice/counselling/incrisis/

Biology Teaching Office:

We are happy to hear from you about teaching matters. The School of Biology Teaching Office is open Monday to Friday 09.00 - 13.00 and 14.00 - 17.00. School of Biology staff will respond to your emails during these hours. Our team will provide a response to you within three working days.

Biology Teaching Office (Level 2), University of St Andrews, Biomolecular Sciences Building, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk

Tel: 01334 46 3602 or 3566

BL3316: Contributing Staff

Dr Verena Dietrich-Bischoff
(Module Organiser)

Lecturer (Education-Focused)vdb@st-andrews.ac.uk
Prof Nathan BaileyProfessornwb3@st-andrews.ac.uk
Dr Miguel BarbosaLecturer in Marine Biologymb334@st-andrews.ac.uk
Dr Verena Dietrich-Bischoff
(Module Organiser)
Lecturer (Education-Focused)vdb@st-andrews.ac.uk
Prof John JonesProfessorjj33@st-andrews.ac.uk
Prof Mike RitchieProfessormgr@st-andrews.ac.uk
Prof Graeme RuxtonProfessorgr41@st-andrews.ac.uk
Dr Harry WatkinsHonorary Research Fellowjhrw@st-andrews.ac.uk
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3316: Learning Outcomes

Students completing module BL3316 successfully should be able to:

  • Explain the basic principles of coevolution at the intra- and interspecific level
  • Understand how coevolution shapes species interactions and can affect ecosystems
  • Compare and contrast detailed examples of coevolutionary interactions in different contexts and systems
  • Discuss the mechanisms that can maintain mutualisms
  • Develop skills for scientific exploration and analysis
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3316: Acquired Skills

Practical Skills

  • Field sampling methods (Botanics)
  • Fixing and preserving specimens
  • Measuring specimens using calipers
  • Species identification (Botany)

Transferable Skills

  • "Short" practical write-up (e.g. completed worksheet)
  • Paper critique
  • Critically evaluating sources/information
  • Finding information on the web
  • Finding literature
  • Referencing
  • Searching databases
  • Data analysis
  • Use Excel
  • Use R or R Studio
  • Lab or field notebook
  • Critiquing experimental design
  • Designing experiments
  • Organising group work
  • Working in pairs/small groups
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Policies

(Information in this section applies to all Biology Modules)

  • The procedures and regulations followed by the School of Biology are outlined in the University Handbook and in the School of Biology UG handbook  JH booklet info (st-andrews.ac.uk) 
  • All coursework associated with the module must be completed and submitted by its due date.
  • Specific School regulations relating to absence reporting, penalties and rules for late submission of work, extensions for coursework, return of coursework, S-coding, Good Academic Practice and Academic Alert are stated in the School of Biology UG hand book JH booklet info (st-andrews.ac.uk)  and students are required to carefully read these regulations.
  • Students are also referred to the University Handbook, available at: http://www.st-andrews.ac.uk/studenthandbook/