Biology Online Module Handbook

BL3307 Evolution

(BL3307 online module handbook version 16)

cover image
Topics in this module will include: molecular variation and evolution, including phylogeny reconstruction; the evolution and maintenance of sex; the genetics of continuous traits, and the relative importance of continuous and discontinuous variation in evolution; evolutionary developmental biology; evolution of population genetic structure; the genetics of speciation, covering the evolution of pre- and post-zygotic isolation, and parapatric, sympatric and island speciation. Practicals will involve computer simulations to investigate a range of evolutionary phenomena, plus use of molecular markers to examine population structure and speciation.*

Credits: 20*

Semester: 1

Module Organiser
Prof Thomas Meagher
trm3@st-andrews.ac.uk
office 3364, lab 3347
Pre-requisite Modules:
BEFORE TAKING THIS MODULE YOU MUST PASS BL2303. BEFORE TAKING THIS MODULE YOU SHOULD PASS BL2303*
Anti-requisite Modules:
None*
Post-requisite Modules:
*
Additional Module Information:
Please check MMS regularly for additional module information


Biology Teaching Office:
Room 1.17, Biomolecular Sciences Building, University of St Andrews, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk      Tel: 01334  463602

BL3307: Timetable

Legend (not all modules have every event type):
lecture
tutorial
workshop
practical
other

Semester 1: Week 1

DATE & TIMEVENUESTAFFEVENT
Monday
18-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Thomas Meagher

Lecture L1: Genetic variation within and between populations
2017-8_BL3307_L1
Tuesday
19-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Thomas Meagher

Lecture L2: Phenotypic Evolution
2017-8_BL3307_L2
Wednesday
20-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Thomas Meagher

Lecture L3: Population Genealogies & Parentage analysis
2017-8_BL3307_L3

Semester 1: Week 2

DATE & TIMEVENUESTAFFEVENT
Monday
25-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Mike Ritchie

Lecture L4: Introduction; Genes and Evolution, a recap and an addition
2017-8_BL3307_L4
Tuesday
26-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Mike Ritchie

Lecture L5: Adaptive Landscapes. The weird world of Twin Peaks
2017-8_BL3307_L5
Wednesday
27-09-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Mike Ritchie

Lecture L6: Genetic Revolutions; selection, drift and gene complexes
2017-8_BL3307_L6

Semester 1: Week 3

DATE & TIMEVENUESTAFFEVENT
Monday
02-10-2017
11:00 to 12:00
Bute Building
Lecture Theater D
Dr Nathan Bailey

Lecture L7: Adaption
2017-8_BL3307_L7
Tuesday
03-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Nathan Bailey

Lecture L8: Contemporary Evolution (Rapid Evolution?)
2017-8_BL3307_L8
Wednesday
04-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Nathan Bailey

Lecture L9: Genomics of Adaptation
2017-8_BL3307_L9

Semester 1: Week 4

DATE & TIMEVENUESTAFFEVENT
Monday
09-10-2017
11:00 to 12:00
Bute Building
Leture Theatre D
Dr Michael Morrissey

Lecture L10: Polygenic inheritance and similarity among relatives
2017-8_BL3307_L10
Tuesday
10-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Michael Morrissey

Lecture L11: Selection and response of continuous traits
2017-8_BL3307_L11
Wednesday
11-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Michael Morrissey

Lecture L12: Genetic correlations and evolutionary constraints
2017-8_BL3307_L12
Friday
13-10-2017
09:00 to 12:00
Irvine Building
Computer Classroom
Dr Michael Morrissey

Practical P1: Playtime: the QG sandbox in R - Group 1
2017-8_BL3307_P1
Friday
13-10-2017
14:00 to 17:00
Bute Building
Computer Classroom
Dr Michael Morrissey

Practical P2: Playtime: the QG sandbox in R - Group 2
2017-8_BL3307_P2

Semester 1: Week 5

DATE & TIMEVENUESTAFFEVENT
Monday
16-10-2017
11:00 to 12:00
Bute Building
Lecture Theater D
Dr Andy Gardner

Lecture L13: Kin selection
2017-8_BL3307_L13
We €™ll look at Price €™s equation of natural selection, and the related concepts of variance, covariance and regression. This will then lead to a general derivation of Hamilton €™s rule of kin selection, and a discussion of the concepts of neighbour-modu
Tuesday
17-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Andy Gardner

Lecture L14: Multilevel Selection
2017-8_BL3307_L14
Returning to Price €™s equation, we €™ll see that this can alternatively be used to underpin a theory of multilevel selection. We €™ll then explore scenarios in which the group can be considered a €˜superorganism €™ in its own right. This will clarify th
Wednesday
18-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Andy Gardner

Lecture L15: Major Transitions in Evolution
2017-8_BL3307_L15
Major transitions in evolution are revolutions in the way that heritable information is stored and transmitted, and often involve irreversible division of labour and the emergence of a new level of individuality. We €™ll explore major transitions from the

Semester 1: Week 7

DATE & TIMEVENUESTAFFEVENT
Monday
30-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Carolin Kosiol

Lecture L16: Phylogenetics and selection I
2017-8_BL3307_L16
Tuesday
31-10-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Carolin Kosiol

Lecture L17: Phylogenetics and selection II
2017-8_BL3307_L17
Wednesday
01-11-2017
11:00 to 12:00
Bute Building
Lecture Theater D
Dr Dave Ferrier

Lecture L18: History and phylogeny
2017-8_BL3307_L18
Friday
03-11-2017
11:00 to 14:00
Bute Building
PC Lab
Prof Thomas Meagher

Practical P3: Hierarchical genetic analysis of population structure - Group 1
2017-8_BL3307_P3
Friday
03-11-2017
14:00 to 17:00
Bute Building
PC Lab
Prof Thomas Meagher

Practical P4: Hierarchical genetic analysis of population structure - Group 2
2017-8_BL3307_P4

Semester 1: Week 8

DATE & TIMEVENUESTAFFEVENT
Monday
06-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Dave Ferrier

Lecture L19: Developmental toolkit
2017-8_BL3307_L19
Tuesday
07-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Dave Ferrier

Lecture L20: Evolution of animal development
2017-8_BL3307_L20
Wednesday
08-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Dave Ferrier

Lecture L21: Palaeontology, problems and prospects
2017-8_BL3307_L21

Semester 1: Week 9

DATE & TIMEVENUESTAFFEVENT
Monday
13-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Anne Magurran

Lecture L22: Community ecology & Darwin
2017-8_BL3307_L22
Tuesday
14-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Prof Anne Magurran

Lecture L23: Ecological and evolutionary change in the contemporary world
2017-8_BL3307_L23
Wednesday
15-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Mike Webster

Lecture L24: Paleoanthropology
2017-8_BL3307_L24

Semester 1: Week 10

DATE & TIMEVENUESTAFFEVENT
Monday
20-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D


Lecture L25: Evolution of symbiotic interactions
2017-8_BL3307_L25
GUEST - Dr Rebecca Yahr, Royal Botanic Garden Edinburgh
Tuesday
21-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Mike Webster

Lecture L26: The genus Homo
2017-8_BL3307_L26
Wednesday
22-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Mike Webster

Lecture L27: Homo sapiens I
2017-8_BL3307_L27

Semester 1: Week 11

DATE & TIMEVENUESTAFFEVENT
Monday
27-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Ildiko Somorjai

Lecture L28: Evolution of Regeneration
2017-8_BL3307_L28
Tuesday
28-11-2017
11:00 to 12:00
Bute Building
Lecture Theatre D
Dr Mike Webster

Lecture L29: Homo sapiens II
2017-8_BL3307_L29
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3307: Reading List

Click for BL3307 reading list

BL3307: Assessment

3-hour Written Examination = 66%, Coursework = 34%*

Assessment:
Practical Report 1
Due by:
18/10/2017 23:59
Feedback due by:
01/11/2017 23:59
Type:
Single Upload
Weight:
20%
Assessment:
Practical Report 2
Due by:
08/11/2017 23:59
Feedback due by:
22/11/2017 23:59
Type:
Single Upload
Weight:
20%
Assessment:
Essay
Due by:
13/11/2017 23:59
Feedback due by:
27/11/2017 23:59
Type:
Single Upload
Weight:
60%

(MMS assessment data cached: 17 November 2017 23:20:14.)


The following related information applies to all Biology modules:

School of Biology Marking Criteria: See School of Biology Undergraduate Handbook
Late submission of continuous assessment work:

All late submissions of coursework that do not require electronic submission should be made via the late submissions box in the Biomolecular Science Building (beside the Teaching Office)
Exam details: See School of Biology Undergraduate Handbook
Exam timetable: see http://www.st-andrews.ac.uk/students/academic/examinations/examtimetablescurrent/
Expected attendance: See School of Biology Undergraduate Handbook for detailed attendance requirements.
Good Academic Practice & Avoiding Academic Misconduct: See School of Biology Undergraduate Handbook
University Student Handbook: University Student Handbook
School and University regulations in the School and University Undergraduate Handbook relating to absence reporting, penalties and rules for late submission of work, extensions for coursework, return of coursework, S-coding, good academic practice and Academic Alerts.: School of Biology Undergraduate Handbook University Student Handbook
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Who to ask

(Information in this section applies to all Biology Modules)

Questions about different aspects of the module should be directed to different people:

Questions about Contact
General teaching matters Biology Teaching Office ( bioteach@st-andrews.ac.uk )
Rescheduled or cancelled events Check your University email
Lecture or practical content The lecturer who presented the material
Completing assessed practical assignments The lecturer who set the assignment
Completing assessments Module Organiser (Prof Thomas Meagher trm3@st-andrews.ac.uk)
Marking on continuous assessment The Demonstrator or Module Organiser (Prof Thomas Meagher trm3@st-andrews.ac.uk)
Marking on exams Module Organiser (Prof Thomas Meagher trm3@st-andrews.ac.uk)
Rearranging practical days Grant Brown
Absence and/or extensions Module Organiser (Prof Thomas Meagher trm3@st-andrews.ac.uk) and the Biology Teaching Office ( bioteach@st-andrews.ac.uk )

Difficulties with academic progress which impact more than one module:

Year Coordinator

See School of Biology Undergraduate student handbook for list: http://biology.st-andrews.ac.uk/documents/UndergraduateHandbook.pdf

Overall performance, progress or future directions:

Advisor of Studies

Disability:

Disability Coordinator (Dr Jacqueline Nairn jn37@st-andrews.ac.uk)

Biology Teaching Office:

Room 1.17, Biomolecular Sciences Building, University of St Andrews, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk

Tel: 01334 463602

BL3307: Contributing Staff

Prof Thomas Meagher

Prof Thomas Meagher
(Module Organiser)

Professortrm3@st-andrews.ac.uk
Dr Nathan BaileyDr Nathan BaileyReadernwb3@st-andrews.ac.uk
Dr Dave FerrierDr Dave FerrierReaderdekf@st-andrews.ac.uk
Dr Andy GardnerDr Andy GardnerReaderag243@st-andrews.ac.uk
Dr Carolin KosiolDr Carolin KosiolLecturerck202@st-andrews.ac.uk
Prof Anne MagurranProf Anne MagurranProfessoraem1@st-andrews.ac.uk
Prof Thomas MeagherProf Thomas Meagher
(Module Organiser)
Professortrm3@st-andrews.ac.uk
Dr Michael MorrisseyDr Michael MorrisseyResearch Fellowmbm5@st-andrews.ac.uk
Prof Mike RitchieProf Mike RitchieProfessormgr@st-andrews.ac.uk
Dr Ildiko  SomorjaiDr Ildiko SomorjaiMASTS Lecturerimls@st-andrews.ac.uk
Dr Mike WebsterDr Mike WebsterLecturer in Behavioural and Evolutionary Biologymmw1@st-andrews.ac.uk
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3307: Learning Outcomes

Students completing module BL3307 successfully should be able to:

  • Understand the basic processes of population genetics underlying evolutionary change.
  • Understand the genetic basis of continuous variation and its importance in population differentiation and speciation.
  • Know how to analyze molecular variation for examining genetic diversity and population structure.
  • Understand how evolution occurs at the molecular level.
  • Develop skills for quantitative analysis of evolutionary change.
  • Acquire experience of working independently and together in groups.
  • Develop skills for scientific exploration and discovery.
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL3307: Acquired Skills

Practical Skills

  • Mendelian genetics

Transferable Skills

  • "Full" practical write-up (Intro, Methods, Results, Discussion)
  • "Short" practical write-up (e.g. completed worksheet)
  • Short essay (1000-2000 words)
  • Computer programming
  • Problem-solving questions
  • Calculations/equations
  • Descriptive statistics
  • Likelihood
  • Non-linear fit
  • Other mathematical models
  • Produce tables
  • Use Excel
  • Use other data analysis software
  • Use R or R Studio
  • Working in pairs/small groups
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Policies

(Information in this section applies to all Biology Modules)