School of Biology current student

Information for current students at the School of Biology

Biology Online Module Handbook

BL2303 Evolutionary Biology

(BL2303 online module handbook version 66)

cover image

image: Nuptial feeding in the dance fly Rhamphomyia. Mating takes place while the female is feeding on a prey item provided by the male, but some males cheat by re-using worthless or fake prey.

Evolution is a fundamentally important component of our understanding of all biological phenomena, from molecular to ecosystem scales. This module will give an overview of the history and major principles of modern evolutionary biology, aimed at contemporary biologists of all backgrounds.*

Credits: 15*

Semester: 1

Module Organiser
Prof Mike Ritchie
mgr@st-andrews.ac.uk
01334 463495
Pre-requisite Modules:
Before taking this module you must pass BL1101 and pass BL1102*
Anti-requisite Modules:
*
Post-requisite Modules:
*
Additional Module Information:


Biology Teaching Office:
Level 2, Biomolecular Sciences Building, University of St Andrews, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk      Phone: +44 (0)1334 46 3602 or 3566

BL2303: Timetable

Legend (not all modules have every event type):
lecture
tutorial
workshop
practical
other

Semester 1: Week 1

DATE & TIMEVENUESTAFFEVENT
Thursday
14-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L1: Course Introduction
2023-4_BL2303_L1
Introduction to the course and a reminder of what Evolution is.
Thursday
14-09-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Prof Mike Ritchie
Dr Kelly Robinson
Practical P1: An introduction to Natural Selection
2023-4_BL2303_P1
Attend on your scheduled day
Friday
15-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L2: Evolution is a fact
2023-4_BL2303_L2
We will review the evidence that compels scientists to conclude that the diversity of life on Earth has arisen due to evolutionary change is more than only a theory.
Friday
15-09-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Prof Mike Ritchie
Dr Kelly Robinson
Practical P2: An introduction to Natural Selection
2023-4_BL2303_P2
Attend on your scheduled day

Semester 1: Week 2

DATE & TIMEVENUESTAFFEVENT
Wednesday
20-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Dr Carolin Kosiol
-
Lecture L3: Phylogenetics1
2023-4_BL2303_L3
An introduction to phylogenetics
Thursday
21-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Dr Carolin Kosiol
-
Lecture L4: Phylogenetics2
2023-4_BL2303_L4
More on Phylogenetics
Friday
22-09-2023
11:00 to 12:00
United College School III
School 1
Dr Carolin Kosiol
-
Lecture L5: Phylogenetics 3
2023-4_BL2303_L5
More on Phylogenetics

Semester 1: Week 3

DATE & TIMEVENUESTAFFEVENT
Thursday
28-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L6: Levels of Selection
2023-4_BL2303_L6
Thursday
28-09-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Dr Carolin Kosiol
-
Practical P3: Phylogenetics
2023-4_BL2303_P3
Attend on your scheduled day
Friday
29-09-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L7: Natural Selection on a gene
2023-4_BL2303_L7
Friday
29-09-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Dr Carolin Kosiol
-
Practical P4: Phylogenetics
2023-4_BL2303_P4
Attend on your scheduled day

Semester 1: Week 4

DATE & TIMEVENUESTAFFEVENT
Wednesday
04-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L8: Natural Selection & Hardy Weinberg
2023-4_BL2303_L8
Thursday
05-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L9: Genetic Drift
2023-4_BL2303_L9
Friday
06-10-2023
11:00 to 12:00
United College School III
School 1
Prof Mike Ritchie
-
Lecture L10: Why have Sex?
2023-4_BL2303_L10

Semester 1: Week 5

DATE & TIMEVENUESTAFFEVENT
Thursday
12-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L11: Sexual reproduction in animals 1
2023-4_BL2303_L11
Darwins second major theory was that sexual selection caused the evolution of elaborate traits that were hard to explain by Natural Selection.
Thursday
12-10-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Prof Mike Ritchie
Dr Kelly Robinson
Practical P5: Bene bags
2023-4_BL2303_P5
Attend on your scheduled day
Friday
13-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L12: Sexual reproduction in animals 2
2023-4_BL2303_L12
How has sexual selection influenced animal evolution?
Friday
13-10-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Prof Mike Ritchie
Dr Kelly Robinson
Practical P6: Bene bags
2023-4_BL2303_P6
Attend on your scheduled day

Semester 1: Week 7

DATE & TIMEVENUESTAFFEVENT
Thursday
26-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L13: Sexual reproduction in plants 1
2023-4_BL2303_L13
Plants show a wide array of breeding systems, and specialisation to either avoid or promote inbreeding are a major driver of breeding system evolution. We will explore this dichotomy leading up to asking whether inbreeding is bad for plants.
Friday
27-10-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L14: Sexual reproduction in plants 2
2023-4_BL2303_L14
Plants exhibit a wide array of gender variation. We will explore evolutionary drivers of gender variation leading up to gender specialisation and separation of male and female function.

Semester 1: Week 8

DATE & TIMEVENUESTAFFEVENT
Wednesday
01-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Nathan Bailey
-
Lecture L15: Mutation
2023-4_BL2303_L15
The process of mutation and its role in evolution
Thursday
02-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Nathan Bailey
-
Lecture L16: Rates of Evolution
2023-4_BL2303_L16
How fast does evolution occur, and does the rate vary?
Thursday
02-11-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Prof Mike Ritchie
Prof Mike Ritchie
Tutorial T1: Hardy Weinberg problems workshop
2023-4_BL2303_T1
Attend on your scheduled day
Friday
03-11-2023
11:00 to 12:00
United College School III
School 1
Prof Nathan Bailey
-
Lecture L17: Fisher and the Geometric model
2023-4_BL2303_L17
What is FGM and why is it important?
Friday
03-11-2023
14:00 to 17:00
Biomedical Sciences Building
Biology Teaching Lab (MBS:142)
Dr Kelly Robinson
Prof Mike Ritchie
Tutorial T2: Hardy Weinberg problems workshop
2023-4_BL2303_T2
Attend on your scheduled day

Semester 1: Week 9

DATE & TIMEVENUESTAFFEVENT
Thursday
09-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Andy Gardner
-
Lecture L18: Another look at Adaptation
2023-4_BL2303_L18
We will explore the logic of adaptation by natural selection, and its application as a predictive, scientific tool.
Friday
10-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Andy Gardner
-
Lecture L19: Social Adaptation
2023-4_BL2303_L19
Social adaptations appear to challenge the notion of the survival of the fittest. We will explore Darwinian musings on kin selection and Hamiltonian theory of inclusive fitness.

Semester 1: Week 10

DATE & TIMEVENUESTAFFEVENT
Wednesday
15-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L20: Species & Biodiversity
2023-4_BL2303_L20
Species diversity and evolutionary potential
Thursday
16-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Dr Kelly Robinson
-
Lecture L21: Reproductive isolation
2023-4_BL2303_L21
What causes reproductive isolation?
Thursday
16-11-2023
14:00 to 17:00
Offsite
St Andrews Botanic Garden
Prof Mike Ritchie
Dr Kelly Robinson
Practical P7: Biodiversity
2023-4_BL2303_P7
Attend on your scheduled day. GUEST James Hearsum, Garden Executive Director, will provide a short introduction.
Friday
17-11-2023
11:00 to 12:00
United College School III
School 1
Prof Mike Ritchie
Prof Mike Ritchie
Lecture L22: Geography & Speciation
2023-4_BL2303_L22
Geographic modes of speciation
Friday
17-11-2023
14:00 to 17:00
Offsite
St Andrews Botanic Garden
Dr Kelly Robinson
Prof Mike Ritchie
Practical P8: Biodiversity
2023-4_BL2303_P8
Attend on your scheduled day. GUEST James Hearsum, Garden Executive Director, will provide a short introduction.

Semester 1: Week 11

DATE & TIMEVENUESTAFFEVENT
Thursday
23-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L23: Evolution and Society
2023-4_BL2303_L23
Evolutionary biology has, since its formalisation as a discipline in the 19th century, generated extraordinary public controversy. What are the benefits of applying evolutionary theory to human societies, and how can this be misused?
Friday
24-11-2023
11:00 to 12:00
Mathematical Institute
LTB
Prof Mike Ritchie
-
Lecture L24: Course conclusion
2023-4_BL2303_L24
We will review what you have learned over the course of the module, and past exam question examples.
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL2303: Assessment

2-hour Written Examination = 50%, Coursework = 50%*

The following related information applies to all Biology modules:

School of Biology Marking Criteria: See JH booklet info (st-andrews.ac.uk) 
Late submission of continuous assessment work:

All late submissions of coursework that do not require electronic submission should be made via the Biology Teaching Office, Level 2, BMS Building, North Haugh.
Exam details: See School of Biology UG Handbook JH booklet info (st-andrews.ac.uk)  : All Biology exams will be conducted online for 2022-23.
Exam timetable: See Timetables - Exams - University of St Andrews (st-andrews.ac.uk) 
Expected attendance: See JH booklet info (st-andrews.ac.uk)  for detailed attendance requirements.
Good Academic Practice & Avoiding Academic Misconduct: See JH booklet info (st-andrews.ac.uk)
University Student Handbook: University Student Handbook
School and University regulations in the School and University Undergraduate Handbook relating to absence reporting, penalties and rules for late submission of work, extensions for coursework, return of coursework, S-coding, good academic practice and Academic Alerts.: JH booklet info (st-andrews.ac.uk)
University Student Handbook
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Who to ask

(Information in this section applies to all Biology Modules)

Before contacting staff, students should check the content of the Biology Undergraduate Handbook, the module handbook and specific task instructions.

Questions about Contact
General teaching matters Biology Teaching Office ( bioteach@st-andrews.ac.uk )
Rescheduled or cancelled events Check your University email
Lecture or practical content The lecturer who presented the material
Completing assessed practical assignments The lecturer who set the assignment
Completing assessments Module Organiser (Prof Mike Ritchie mgr@st-andrews.ac.uk)
Marking on continuous assessment The Demonstrator or Module Organiser (Prof Mike Ritchie mgr@st-andrews.ac.uk)
Marking on exams Module Organiser (Prof Mike Ritchie mgr@st-andrews.ac.uk)
Rearranging practical days
Module Organiser (Prof Mike Ritchie mgr@st-andrews.ac.uk)
Absence and/or extensions Module Organiser (Prof Mike Ritchie mgr@st-andrews.ac.uk)

and the Biology Teaching Office ( bioteach@st-andrews.ac.uk )

Difficulties with academic progress which impact more than one module:

Year Coordinator

See School of Biology UG Handbook for list:
JH booklet info (st-andrews.ac.uk)

Overall performance, progress or future directions:

Advisor of Studies

Disability:

Disability Coordinator ( biodisabilities@st-andrews.ac.uk )

For advice and support on any issue e.g. academic, financial, international, personal or health matters, or if you are unsure of who to go to for help:

Advice & Support Centre

Address: 79 North Street, St Andrews

Email: theasc@st-andrews.ac.uk

Web: https://www.standrews.ac.uk/ask-a-question/

Tel: 01334 462020

University assistance with urgent matters out of office hours:

Tel: 01334 476161

Web: https://www.st-andrews.ac.uk/students/advice/counselling/incrisis/

Biology Teaching Office:

We are happy to hear from you about teaching matters. The School of Biology Teaching Office is open Monday to Friday 09.00 - 13.00 and 14.00 - 17.00. School of Biology staff will respond to your emails during these hours. Our team will provide a response to you within three working days.

Biology Teaching Office (Level 2), University of St Andrews, Biomolecular Sciences Building, North Haugh, St Andrews, Fife KY16 9ST

Email: bioteach@st-andrews.ac.uk

Tel: 01334 46 3602 or 3566

BL2303: Contributing Staff

Prof Mike Ritchie
(Module Organiser)

Professormgr@st-andrews.ac.uk
Prof Nathan BaileyProfessornwb3@st-andrews.ac.uk
Prof Andy GardnerProfessor of Biologyag243@st-andrews.ac.uk
Dr Carolin KosiolReaderck202@st-andrews.ac.uk
Prof Mike Ritchie
(Module Organiser)
Professormgr@st-andrews.ac.uk
Dr Kelly RobinsonAssociate Lecturer in Animal Behaviour and Evolutionkjr33@st-andrews.ac.uk
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL2303: Learning Outcomes

Students completing module BL2303 successfully should be able to:

  • Develop an understanding of Evolution and its history
  • Understand Natural Selection and Genetic Drift
  • Develop knowledge of Molecular Evolution and mutation
  • Understand the evolution and consequences of sexual reproduction
  • Know the relationship between speciation and Biodiversity
  • Understand the basic principles of Phylogenetics
  • Appreciate the role of the study of evolution to contemporary society
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

BL2303: Acquired Skills

Practical Skills

  • Field sampling methods (Botanics)
  • Mendelian genetics

Transferable Skills

  • "Full" practical write-up (Intro, Methods, Results, Discussion)
  • Short essay (1000-2000 words)
  • Critically evaluating sources/information
  • Finding information on the web
  • Finding literature
  • Referencing
  • Searching databases
  • Lab safety awareness
  • Problem-solving questions
  • Biodiversity analysis
  • Calculations/equations
  • Chi-square test
  • Data analysis
  • Data presentation
  • Descriptive statistics
  • Genetic Drift
  • Phylogenetic analysis
  • Produce graphs/figures
  • Produce tables
  • Use other data analysis software
  • Lab or field notebook
  • Critiquing experimental design
  • Generate class dataset
  • Working in pairs/small groups
Data Sources: information in this online module handbook is gathered from several sources.
Small footnote marks beside content, or by section titles indicate the source.
* University of St Andrews Module Catalogue
School of Biology Symbiosis Database (module organisers only)
University of St Andrews MMS (St Andrews users only)

Policies

(Information in this section applies to all Biology Modules)

  • The procedures and regulations followed by the School of Biology are outlined in the University Handbook and in the School of Biology UG handbook  JH booklet info (st-andrews.ac.uk) 
  • All coursework associated with the module must be completed and submitted by its due date.
  • Specific School regulations relating to absence reporting, penalties and rules for late submission of work, extensions for coursework, return of coursework, S-coding, Good Academic Practice and Academic Alert are stated in the School of Biology UG hand book JH booklet info (st-andrews.ac.uk)  and students are required to carefully read these regulations.
  • Students are also referred to the University Handbook, available at: http://www.st-andrews.ac.uk/studenthandbook/